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She retrieves three survivors — the British nurse Claire Paige Olivia Williams and two men, one of them wounded — from the British hospital ship Fort James, which had been sunk two days earlier; one of the survivors blames the sinking on a German U-boat that he briefly saw on the surface just before Fort James suffered a torpedo hit.
As they pick up the survivors, the crew of Tiger Shark spots a German destroyer bearing down on them. The submarine has several encounters with the German destroyer and suffers damage from depth charges in the process. Brice shoots Schillings dead when the German panics and grabs a scalpel to defend himself. Brice reveals to Paige that Tiger Shark had sunk a German submarine tender recently and that her previous commanding officer, Lieutenant Commander Winters Nick Hobbs , had died after Tiger Shark surfaced to confirm the sinking.
According to Brice, Winters was using a boathook to try to obtain a souvenir from the flotsam left behind by the sunken German ship when Tiger Shark struck a submerged object, causing Winters to hit his head while reaching for his souvenir, fall overboard, and drown before he could be rescued.
Brice then assumed command of Tiger Shark. Soon after the death of the German prisoner, those aboard Tiger Shark begin to hear disembodied voices and experience other eerie events. Winters, on deck with only Brice, Coors, and Lieutenant Paul Loomis Holt McCallany , had ordered them to have a gunnery party come on deck to machine-gun the survivors of the sunken German ship in the water.
When Brice, Loomis, and Coors objected, a heated argument had broken out and escalated into a physical altercation during which Winters hit his head and fell overboard. Before they leave the ballast tank, Coors dies in a mysterious accident. He escapes from the submarine via an escape trunk while the submarine is underwater, and dies when he is impaled on an outside railing.
A series of bizarre mechanical problems causes the crew of Tiger Shark to lose control of the submarine, and the submarine turns back towards the site of her sinking of the German ship, apparently of her own volition.
Meanwhile, crewmen die in accidents at an alarming rate. Paige and Odell discover that Tiger Shark mistook Fort James for the German submarine tender and sank Fort James instead; they also learn that Brice, Loomis, and Coors believed that they could not afford this drastic mistake to appear on their records and conspired to suppress the story, killing Winters on the deck of Tiger Shark as he tried to save the survivors of Fort James. Tiger Shark ultimately is crippled by a mounting number of accidents, and only five of those aboard remain alive: Wally concludes that Tiger Shark is haunted by a "malediction" that must be satisfied in order to escape its netherworld between heaven and hell.
After Tiger Shark arrives at the location of the sinking of Fort James, she surfaces in a disabled condition and those aboard her detect a surface ship nearby. Brice prevents the surviving crew of Tiger Shark from radioing the nearby ship, but Paige sneaks out on deck and tries to signal the ship with a flashlight.
Brice confronts her and holds her at gunpoint. His remorse over the accident overcomes him; he admits the entire cover-up to Paige and then shoots himself in the head, falling dead into the ocean.
The ship Paige is signaling turns out to be British and picks up the four survivors of Tiger Shark. As they look on, Tiger Shark sinks, coming to rest on the ocean floor next to the wreck of Fort James.